Family Tenebrionidae

(+ Lagriidae and Alleculidae)

The darkling beetles

Reviewed, corrected, and supplemented by C.A. Triplehorn, May 1993, May 1995; R. Aalbu, June 1993; J. Doyen, March 1994; W.E. Steiner Jr., January 1997.

The higher classification and generic placement used here follows that of Aalbu (ms 1992).

Division Lagriiformes

Subfamily Lagriinae

The long-jointed bark beetles

Tribe Statirini

Statira Serville (see Parsons 1975)

• S. basalis Horn, TX-MO-NC, FL (STJ)
• S. croceicollis Maeklin, MS-MD, FL (DAD, MON, OSC, PAL, PIN, VOL)
• S. dolera Parsons, GA-FL (ALA, BRE, FLA, MAN, MON, PAL, PUT, VOL)
• S. gagatina Melsheimer, eUS, FL?
• S. liebecki Leng, AL-FL (HIG, OKE, PIN, PUT, VOL)

Tribe Goniaderini

Anaedus Blanchard

• A. brunneus (Ziegler), eUS, FL (ALA, HIG)

Paratenetus Spinola

• P. punctatus Spinola, FL (ALA, BRE, IND, LAK, LEV, MARI, PUT, SRO), CAmer

Tribe Lupropini

Prateus LeConte

• P. fusculus LeConte, eUS, FL

Subfamily Phrenapatinae

Tribe Penetini

Dioedus LeConte

• D. punctatus LeConte, eUS, FL (ALA, MARI), Puerto Rico

Division Pimeliiformes

Subfamily Pimeliinae

Tribe Epitragini

(see Freude 1967)

Bothrotes Casey

• B. canaliculatus acutus (LeConte), TX, FL (ALA, IND, LAK, MARI, POL, VOL)
  = fortis Casey
• B. canaliculatus arundinis (LeConte), eUS, FL

Epitragodes Casey

• E. t. tomentosus (LeConte), GA, FL (ALA, DAD, CHA, COLL, IND, LAK, LEE, LEV, POL), Bahamas
  = floridanus Casey, FL
  = obesulus Casey, FL
• E. t. macilentus Casey, AL, GA, NC, FL (ALA, BAK, COLU, GUL, HAR, HIG, MARI, MART, ORA, TAY)
  = cuprascens Casey, FL
  = debilicollis Casey, FL
  = pardalis Casey, FL

Schoenicus LeConte

• S. puberulus LeConte, NJ-FL (ALA, CLA, BRO, COLL, GAD, IND, LIB, MAN, PAL, SRO)

Tribe Trientomini

Trientoma Solier

• *T. wickhami Casey, FL?, Bahamas

Tribe Coniontini

Branchus LeConte

• B. floridanus LeConte, FL (DAD, MON) (see Triplehorn and Weems 1964)

Division Tenebrioformes

Diaperine Lineage

Subfamily Diaperinae

Tribe Phalerini

Phaleria Latreille (see Triplehorn and Watrous1979)

• P. picipes Say, MD-FL (DAD, BRO, IND, MART, MON, NAS, PAL, SJO), Bahamas, Greater Antilles, Lesser Antilles, Mexico, CAmer; in coastal sand beaches
• P. punctipes LeConte, FL (BRO, DAD, DUV, FLA, IND, MON, PAL, PIN, SAR, VOL), Cuba, Jamaica, Bahamas, Lesser Antilles; in coastal sand beaches
• P. testacea Say, ME-FL (DAD, ESC, FLA, FRA, IND, MART, MON, NAS, SJO, VOL), Bahamas, Lesser Antilles, Mexico, Brazil, Argentina; in coastal sand beaches
  = longula LeConte

Tribe Rhipidandrini

Rhipidandrus LeConte
= Cherostus Waterhouse

• R. fulvomaculatus (Dury), FL (HIG, LEV)
• R. paradoxus (Palisot), eUS, FL (ALA, HIG, LAK, LEV, LIB, MARI, SRO)
  = R. flabellicornis (Stürm)

Tribe Bolitophagini

Bolitophagus Illiger

• B. corticola Say, eUS, FL (ALA, LIB, OKA)

Bolitotherus Candèze

• B. cornutus (Panzer), eUS, FL (ALA, COLU, HIG, HIL, LAK, MARI, PUT, UNI)

Tribe Diaperini

Subtribe Adelinina

Adelina Dejean
= Doliema Pascoe

• A. bidens (Schaeffer), FL (BRO, DAD, HIG, IND, MON), Cuba, Hispaniola
• A. pallida (Say), FL (ALA, DAD, DIX, ESC, HIG, PUT, SEM), WInd, Mexico, CAmer, SAmer
• A. plana (Olivier), FL (DAD, HIG, IND, MARI, MON, POL), WInd, Mexico, CAmer, SAmer

Gnatocerus Thunberg

• G. cornutus (Fabricius), the broadhorned flour beetle, cosmopolitan, US, FL (ALA, IND, LEO), Cuba, Mexico to Brazil; a stored products pest
• G. curvicornis (Champion), FL (MON), Mexico, SAmer
• G. guatemalensis (Champion), eUS, FL (ALA, HIG, IND, LEO, MARI, MON), Cuba, Mexico, Guatemala
• G. maxillosus (Fabricius), the slenderhorned flour beetle, cosmopolitan, US, FL (ALA, ESC, GAD, LIB, MARI, MON, SEM), WInd, SAmer

Sitophagus Mulsant

• S. hololeptoides (Laporte), CA, AZ, TX, FL (ALA, COLU, GAD, HIL, LIB, LEO, MAD, MARI), WInd, CAmer, Samer; in stored products

Subtribe Diaperina

Diaperis Geoffroy (see Triplehorn, 1965)

• D. maculata Oliver, eUS, FL (ALA, BRE, CHA, DAD, DUV, GAD, HER, HIG, IND, JEF, LEO, LEV, LIB, MARI, MON, ORA, PAL, PIN, TAY, VOL), CAmer, widespread WInd
  = floridana Blatchley, FL
• D. nigronotata Pic, TX-IA-SC-FL (ALA, CLA, COLU, LIB, PIN, PUT, UNI)
  = bicoloriceps Pic

Liodema Horn

• L. laeve (Haldeman), NC-FL (ALA, DIX, HER, MARI, PUT)

Neomida Latreille
= Hoplocephala Laporte and Brullé

• N. bicornis (Fabricius), TX-MN-MA-FL (ALA, DAD, HER, MARI, MON), WInd
  = viridipennis Fabricius
• N. ferruginea (LeConte), TX-AL-FL (ALA, DAD, HIG, IND, MARI, PIN, POL, VOL), Cuba, Haiti, Mexico, CAmer

Ulomoides Blackburn
= Martianus Fairmaire
= Palembus Casey

• +U. ocularis (Casey), FL (DAD, MON), Puerto Rico, Jamaica, Bahamas, Montserrat; immigrant from Australian Region; in tamarind pods (see Triplehorn and Weems1964)

Platydema Laporte and Brullé

• P. cyanescens Laporte and Brullé, TX-TN-FL (ALA, BRO, DAD, DIX, COLU, HIG, IND, LAK, LEV, MARI, MON, PIN, PUT)
• P. ellipticum (Fabricius), eUS, FL (ALA, BRO, COLU, DAD, DIX, DUV, IND, LAF, LAK, LEO, LEV, LIB, MARI, NAS, ORA, PUT)
• P. erythrocerum Laporte and Brullé, LA, FL (ALA, BRO, DAD, HIG, LEV, LIB, MARI, UNI), SAmer
• P. excavatum (Say), eUS, FL (ALA, BRO, COLU, DAD, LIB, MON, PUT), Mexico, WInd, CAmer, SAmer
• P. flavipes (Fabricius), eUS, FL (ALA, BRO, CLA, COLU, DAD, DUV, HIG, IND, LAK, LEV, LIB, MARI, ORA, POL, PUT, SRO)
• P. laevipes Haldeman, IN, FL (ALA, COLL, LEV, LIB, MARI, PUT, WAK)
• P. micans Horn, seUS, FL (ALA, BRE, BRO, CLA, COLL, DAD, ESC, FLAG, HIG, IND, LEV, LIB, MARI, MON, PAL, SJO, SRO, VOL), Cuba, Jamaica, Haiti, Mexico, Brazil
• P. nigratum (Motschulsky), AZ-TX-AL-FL (BRO, DAD, HIG, IND, MON, SAR), WInd, Costa Rica, Mexico
  = subquadcatum Motschulsky
• P. picilabrum Melsheimer, FL (DIX, MARI, NAS, PUT), IN
• P. ruficolle Laporte and Brullé, eUS, FL (ALA, DIX, ESC, FLA, HIG, IND, LEV, LIB, MARI)
• P. ruficorne (Sturm), eUS, FL (ALA, BAK, CLA, COLU, DAD, DUV, GAD, IND, JAC, LAF, LAK, LIB, MARI, MON, NAS, PUT, SAR)
• P. subcostatum Laporte and Brullé, eUS, FL (ALA, GAD, JAC, LAF, LEV, LIB, MARI, OKA)
• P. n.sp., FL (PAL), Panama

Tribe Crypticini

Gondwanocrypticus Español
= Crypticus Latreille

• G. obsoletus (Say), seUS, FL (ALA, DAD, FRA, HAM, HIL, HIG, IND, LEV, POL, MARI, MON)

Poecilocrypticus Gebien

• +P. formicophilus Gebien, FL (ALA, LEO, LEV, LIB); found in 1978, immigrant from SAmer (see Steiner 1982)

Tribe Trachyscelini

Trachyscelis Latreille

• T. flavipes Melsheimer, Gulf States, NC, FL (DAD), Antilles

Subfamily Hypophloeinae

Tribe Hypophloeini

Corticeus Piller and Mitterpacher (see Triplehorn 1990)

• C. glaber (LeConte), TX-OH-NY-FL (ALA, CHA, DAD, LIB)
• C. parallelus (Melsheimer), TX-NB-PQ-MA-FL (ALA, DUV, LEE, LIB, NAS)
• C. thoracicus (Melsheimer), eNAmer, FL (ALA, COLL, DAD, IND, LIB, MARI, MON, PUT)
  = piliger (LeConte)
• C. tensicollis Triplehorn, FL (BRO, DAD), Cuba, Puerto Rico, Hispaniola (see Triplehorn 1979)

Subfamily Opatrinae

Tribe Opatrini

Blapstinus Latreille (based on unpublished Ph.D. thesis of J.C. Davis)

• B. aciculus Blatchley, FL (DAD, IND, MON), Bahamas; on dunes above beach
• B. alutaceus Casey, FL endemic (MON); on dunes above beach
  = opacus LeConte
• B. dispar Casey, FL endemic (BRO, DAD, PAL); on dunes above beach
• B. fortis Le Conte, TX, FL (ALA, BRE, DAD, FLA, HEN, HIG, HIL, IND, LEV, MARI, MON, OKE, ORA, PAL, PIN, SAR), Cuba, Bahamas, Mexico, CAmer, Hawaii
• B. fuscus Casey, TX, FL (DAD, MON), Mexico
  = humilis Casey
• B. histricus Casey, AZ, FL (BRO, DAD, in V. Golia Coll.); immigrant from southwestern US, possibly on shipments of date palms
• B. metallicus (Fabricius), Can-FL (ALA, BAY, CHA, DAD, FLAG, HIG, HIL, IND, JEF, LEE, LEO, LEV, MARI, PUT, SEM, SRO)
  = Blapstinus aeneolus Melsheimer
  = Blapstinus interruptus LeConte
  = Opatrinus interruptus Say

Cybotus Casey

• C. estriatus (LeConte), FL (BRE, BRO, DAD, PAL, SLU), Mexico; on dunes above beach

Idiobates Casey

• I. castaneus (Knoch), eUS, FL (ALA, LEO, PAL)

Pseudephalus Casey

• P. brevicornis Casey, AL, FL (BAY, ESC, FRA, OKA, SRO, WAK, WAL), Honduras, Costa Rica; on dunes above beach

Ulus Horn

• U. maritimus Casey, TX, FL (BAY, FRA, LEV, OKA, WAL); on dunes above beach

Tribe Platynotini

Opatrinus Latreille

• O. minimus (Palisot de Beauvois), eUS, FL (ALA, COLL, DAD, DIX, GLA, HIG, IND, LAK, MON, OKA, WAK)
• O. pullus (Sahlberg), FL (MON), Mexico, Camer, WInd

Tribe Leichenini

Leichenum Blanchard

• +L. canaliculatum variegatum (Klug), AL-FL (ALA, BRO, CLA, DAD, DIX, DUV, HAR, HIG, IND, LAK, LEE, LEV, MARI, ORA, PAL, POL, SLU, SRO; probably throughout state); immigrant from Madagascar (Woodruff 1963); especially on sandy soil

Tribe Ulomini

Eutochia LeConte

• E. picea (Melsheimer), TX, eUS, FL (ALA, ESC, FLA, GAD, HIG, LAK, LIB, MARI) (According to Steiner, more than one species may be represented here)

Hypogena Dejean

• H. tricornis (Dalman), TX, FL (DAD, IND, STL), Mexico

Uloma Laporte

• U. imberbis LeConte, eUS, FL (ALA, BAY, BRA, DIX, JAC, MARI, PUT)
• U. mentalis Horn, TX-KS-FL (ALA, LEV, LIB, PUT)
• U. punctulata LeConte, eUS, FL (ALA, BAK, COLL, DAD, DUV, HIG, IND, JAC, LAF, LEO, LEV, LIB, MARI, PAL, POL, PUT, SRO, Lake Pickett)

Tribe Helopini

Helops Fabricius

• H. cisteloides Germar, seUS, FL (LIB)
• H. n.sp. 1, FL (n)
• H. n.sp. 2, FL (n)

Nautes Pascoe

• N. viridimicans Horn, FL (BRO, CIT, COLL, DAD, HAR, LAK, LEV, MON, SAR, SLU, VOL)

Tarpela Bates

• T. undulata (LeConte), eUS, FL

Subfamily Tenebrioninae

Tribe Triboliini

Latheticus Waterhouse

• +L. oryzae Waterhouse, cosmopolitan, FL (ALA, LIB, MAD, MARI); a stored products pest
• +L. prosopis Chittenden, cosmopolitan, FL

Lyphia Mulsant and Rey

• +L. tetraphylla Fairmaire, cosmopolitan, FL (LIB)

Palorus Mulsant

• +P. subdepressus (Wollaston), the depressed flour beetle, FL (ALA, MARI), cosmopolitan; a stored products pest
• +P. ratzeburgi (Wissmann), the smalleyed flour beetle, eUS, FL, Old World, cosmopolitan; a stored products pest
• +P. foveicollis Blair, FL (ALA), cosmopolitan; a stored products pest

Tharsus LeConte

• T. seditiosus LeConte, seUS, FL (ALA, DAD, IND, SAR)

Tribolium MacLeay

• +T. audax Halstead, the American black flour beetle, cosmopolitan, FL; in stored products
• +T. castaneum (Herbst), the red flour beetle, US, FL (ALA, COLU, DAD, DUV, HIL, IND, MARI, MON, SEM), WInd, SAmer, cosmopolitan; in a great many kinds of stored products
• +T. confusum Jacquelin du Val, the confused flour beetle, US, FL (ALA, DAD, ORA), WInd, SAmer, cosmopolitan; in a great many kinds of stored products

Tribe Alphitobiini

Alphitobius Stephens

• +A. diaperinus (Panzer), the lesser mealworm beetle, eUS, FL (ALA, BRO, CHA, CLA, DAD, HIL, IND, MAN, MARI, ORA, PAS, PIN, POL, PUT, VOL), Cuba to Puerto Rico, Old World; a pest in many kinds of dried materials
• +A. laevigatus (Fabricius), the black fungus beetle, eUS, FL (BRO, DAD, HIG, IND, MARI, SEM), Cuba, WInd, CAmer, SAmer; a pest in many kinds of dried materials
  = piceus (Olivier)

Tribe Tenebrionini

Neatus LeConte

• N. tenebrioides Beauvois, eUS, FL (LEV, LIB, PUT)

Tenebrio Linnaeus

• +T. molitor Linnaeus, the yellow mealworm beetle, FL (ALA, JEF), cosmopolitan; in many kinds of stored products
• +T. obscurus Fabricius, the dark mealworm beetle, FL?, cosmopolitan; in stored products

Zophobas Blanchard (see Tschinkel 1984)

• Z. atratus (Fabricius), CA-FL (MON), Cuba, SAmer (There is a species of Zophobas commonly sold in Florida pet stores as the "giant mealworm," and has been called in pet publications "Z. morio". It is not conspecific with the species listed here.)
  = morio (Fabricius)
  = rugipes Kirsch

Tribe Amarygmini

Cymatothes Solier
= Pyanisia Laporte

• C. opaca (Solier), seUS, FL (DAD, MON), Mexico
• *C. tristis (Laporte), AL, Cuba, Puerto Rico, Mexico to Panama

Meracantha Kirby

• M. contracta (Beauvois), eUS, FL (LIB)

Subfamily Alleculinae

The comb-clawed beetles

reviewed and corrected by J. M. Campbell April 1993

Tribe Alleculini

Andrimus Casey

• A. murrayi LeConte, FL endemic (ALA, CLA, DAD, HER, HIG, IND, LAK, LEV, LIB, MARI, MON, PUT, SAN, SUW,UNI, VOL)
  = brunneus Casey, FL
  = confusus Casey, FL
  = nigresceus Casey, FL
  = parvulus Casey, FL

Androchirus LeConte

• A. femoralis Olivier, seUS, FL (ALA, COL, LIB, PUT, UNI)

Charisius Champion

• (C. interstitialis Champion, not in Florida)
  = floridanus Linell

Hymenochara Campbell

• H. rufipes (LeConte), MI-NY-FL (MARI)

Hymenorus Mulsant

• H. arkansanus Fall, AR-FL (DAD, LIB)
• H. caducus Fall, AL-FL (ESC)
• H. convexus Casey, (TX?), FL (MON), Cuba, Bahamas
• H. densus LeConte, eUS, AL-FL (DAD, FLA, HIG, MON, WAK), Bahamas, Mexico
• H. dichrous Blatchley, FL endemic (ALA, BRO, HER, SAR)
• H. distinctus Fall, seUS, FL (MARI, POL)
• H. dorsalis Schwarz, FL endemic (COLL, DAD, HIG, IND)
  = sabalensis Blatchley
• H. dubius Fall, MS-AL-FL (ALA, COLL, DIX, LAK, LEO, LEV, LIB, MON, PUT, SAN)
• H. floridanus Casey, FL endemic (IND, MON, ORA)
• H. fuscipennis Fall, FL endemic (HIG, MARI)
• H. granulatus Blatchley, FL endemic (ALA, IND)
• H. heteropygus Fall, FL endemic (ALA, FLA, LEV, LIB)
• H. humeralis LeConte, KY-FL (ALA, LIB)
• H. illusus Fall, AL-FL (ALA, DAD, LIB)
• H. inopiatus Fall, GA-FL (LIB, MON)
• H. melsheimeri Casey, MI-IN, FL (ALA, IND, LEV)
• H. niger Melsheimer, eUS, FL (ALA, COLL, DAD, HILL, IND, LAK, MARI, MON)
• H. obesus Casey FL (FLA, LEO, LIB)
• H. quietus Fall, MO-FL (MON)
• H. semirufus Fall, FL endemic (HIG, IND)
• H. sobrinus Casey, FL endemic (HIG, MAN)
• H. tenellus Casey, FL endemic (ALA, GUL, PUT, SAN)
  = elbertae Blatchley
• H. tenuistriatus Fall, AL-NC-FL (MARI)

Isomira Mulsant

• *I. angusta (Casey), GA
• I. iowensis Casey, IA-FL (ALA, DUV, LIB, MAD, MARI)
• I. oblongula Casey, IN-NY-FL (ALA, DIX)
• I. pulla Melsheimer, RI-NC-FL (ALA, LIB, SAR, UNI)
  = ignova Blatchley
• I. sericea Say, eUS, FL (LIB)
• I. valida Schwarz, FL endemic (ALA, LIB)

Lobopoda Solier

• L. erythrocnemis Germar, seUS, FL (ALA, BAK, BRO, COL, DAD, DIX, HIG, HIL, IND, LAK, LEV, LIB, MON, ORA, OSC, PIN, PUT, SLU, VOL)
• L. nigrans (Melsheimer), eUS, FL (DAD, DUV, ESC, HIG, IND, LAK, PIN)
  = atra Say, eUS, FL
• L. opacicollis Champion, TX-LA-FL (MON), Mexico, CAmer
• L. punctulata Melsheimer, eUS, TX-FL (ALA, DAD, DIX, DUV, HIG, JAC, LEO, LIB, MAN, MON, OKA), Mexico
• L. socia (LeConte), TX-LA-FL (ALA, BRE, BRO, DAD, ESC, HIL, MARI, ORA, PIN, SLU, VOL), Nicaragua; (see Campbell 1986)

Mycetochara Berthold

• M. haldemani LeConte, eUS, FL
• (M. puncticollis Blatchley, is a melandryid (Campbell 1977))

Onychomira Campbell (see Campbell 1984)

• O. floridensis Campbell, FL endemic (HIG)

Subfamily Coelometopinae

Tribe Coelometopini

Alobates Motschulsky

• A. morio (Fabricius), eUS, FL (ALA, BAK, BRO, COLL, DAD, GULF, HIG, IND, LAF, LAK, LEV, MARI, MON, OKE, PUT, SAR)
  = A. barbata Knoch
• A. pennsylvanicus (Degeer), eUS, FL (ALA, BRO, CLA, COLU, DAD, DUV, GIL, GULF, HIG, JAC, LAF, LAK, LEV, LIB, MARI, MON, OKA, ORA, PUT, SAR, SLU, SRO, VOL), Mexico

Centronopus Solier

• C. calcaratus (Fabricius), eUS, FL (ALA, LIB)

Glyptotus LeConte

• G. cribratus LeConte, TX, FL (ALA, BAY, BRO, CLA, DAD, DES, DIX, DUV, HIG, IND, LAK, LEV, MARI, MON, OKA, PUT), Bahamas

Haplandrus LeConte

• H. ater (LeConte), seUS, FL (ALA, CIT, COLU, DUV, FLA, LEV, LIB, MAN, MARI, OKA, PUT, SRO, WAK); rarely collected, but seems to be most abundant in leaf nests of squirrels
• H. collaris Casey, FL endemic (a synonym of H. ater?)
• H. fulvipes (Herbst), eUS, FL (ALA)

Merinus LeConte

• M. laevis (Olivier), Ont-OK-FL (ALA, HAM, JAC)

Polypleurus Eschscholtz

• P. geminatus Solier, seUS, FL (ALA, BAK, BRA, CIT, CLA, HER, HIG, LAK, LEO, LEV, MARI, OKA, ORA, OSC, POL, PUT, SRO, SUM, WAL)
• P. perforatus (Germar), seUS, FL (ESC, JAC, LIB)

Polopinus Casey (see Kritsky 1989)

• P. hubbelli Kritsky, FL endemic (MON)
• P. ingens Casey, GA, FL (ALA, BAK, BRA, CLA, DUV, FRA, GIL, LAK, LEE, LEO, LEV, LIB, MARI, NAS, PIN, POL, PUT, SUW)
• P. nitidus (LeConte), FL endemic (ALA, DIX, HIG, LAK, LIB, MARI, OSC, PAS, POL, PUT, VOL)
  = brevior Casey
  = subdepressus Casey
• P. youngi Kritsky, FL endemic (BRO, COLL, DAD, HAR, GIL, IND, LEE, MARI, MART, MON, OSC, PAL, POL, SRO, SEM)

Xylopinus LeConte

• X. saperdioides (Oliver), eUS, FL (ALA, CIT, COLU, DIX, HIG, IND, LEV, LIB, MARI, OKA)
  = rufipes (Say), a red legged phase or race

Tribe Talanini

Talanus Mäklin (see Triplehorn 1968)
= Dignamptus LeConte

• T. langurinus (LeConte), TX, FL (BRE, BRO, COLL, DAD, GLA, HEN, MON, PAL, SEM, VOL)
• T. spilmani Triplehorn, FL (BRO, COLL, DAD, GLA, MON, OKE, PAL, PUT)
• T. stenochinus (LeConte), LA, FL (ALA, BRE, BRO, DAD, DUV, GLA, HIG, IND, LEE, LEV, MON, PAL, PIN)
  = okeechobeensis Blatchley
• T. sp., FL (HIG, in V. Golia Coll.); possible immigrant from Neotropics

Tribe Strongyliini

Strongylium Kirby (see Triplehorn and Spilman 1973)

• S. anthrax Schwarz, FL endemic (ALA, DAD, GLA, IND, MARI, PAS, PUT)
• S. crenatum Maeklin, TX-OH-FL (ALA, LIB, MARI, POL)
• S. simplicicolle LeConte, MS-NC-FL (ALA, BRA, BRO, COLL, HIG, POL, SAR)
• S. tenuicolle (Say), OK-ON-FL (JAC, LIB, MARI, VOL, WAK)
• S. terminatum (Say), OK-NY-FL

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