Family Dytiscidae

The predaceous water beetles

reviewed and supplemented by R. Roughley, August 1992; checked by J.H. Epler, December 1995

Subfamily Copelatinae

Agaporomorphus Young

• A. dodgei Young, FL endemic (ALA, BAK, COLU, DAD, MAD, MARI, MART, PUT)

Copelatus Erichson

• C. blatchleyi Young, FL endemic (MON)
• C. caelatipennis princeps X angustatus, FL (DAD, PAL)
• C. caelatipennis princeps Young, IL, NJ-LA, FL (ALA, BRE, BRO, CAL, CHA, CLA, COLL, COLU, DAD, DIX, DUV, ESC, FRA, HEN, HIG, HIL, JAC, LEE, LEO, LEV, LIB, MON, NAS, OKA, OKE, ORA, OSC, PAL, PIN, POL, PUT, SJO, SRO, SAR, SEM, TAY, VOL, WAK), Bahamas (Andros)
• C. chevrolati chevrolati Aubé, MI?, IL-TX, FL (ALA, BRO, COLU, DAD, DUV, HEN, MAD, MON, OSC, PIN, SJO, SRO, TAY, VOL), Bahamas (Andros)
• C. cubaensis Schaeffer, FL (DAD), Cuba, Guadeloupe, CAmer
• C. glyphicus (Say), TX, CA-NS, FL (ALA, BAY, COLU, GLA, HAM, LIB, POL)

Subfamily Laccophilinae

Agabetes Crotch

• A. acuductus (Harris), ON, MA-AR, FL (ALA, GAD, WAK)

Laccodytes Régimbart

• L. pumilio (LeConte), FL (ALA, DAD, HIG, POL, VOL), Cuba

Laccophilus Leach

• L. fasciatus rufus Melsheimer, FL (ALA, DUV, GAD, ESC, JAC, LIB)
• L. gentilis gentilis LeConte, FL (ALA, BAK, BRO, DAD, DUV, FLAG, GLA, HEN, HER, HIG, LEE, LEV, OKE, ORA, PAL, PIN, SAR, TAY, VOL), ?Cuba
• L. proximus Say, FL (ALA, BAY, BRE, BRO, CHA, COLL, DAD, DES, DUV, FRA, GAD, GUL, HEN, HER, HIG, HIL, LEE, LEO, MART, OKE, ORA, PAL, PIN, POL, PUT, SJO, SAR, SEM, TAY, VOL, WAK), Bahamas, Cuba, Jamaica, Haiti, Puerto Rico, Barbados, Virgin Islands
• L. vacaensis vacaensis Young, AZ-LA, FL (MON), Bahamas, Cuba, Haiti, Mexico, Guatemala

Subfamily Hydroporinae

Tribe Methlini

Celina Aubé

• C. angustata Aubé, TX-MA-FL (ALA, BRO, DAD, DUV, HIL, LEE, OKA, PUT, SJO, VOL)
• C. contiger Guignot, TN, GA-FL (PIN)
• C. grossula LeConte, LA-GA-FL (ALA, HIG)
• C. hubbelli Young, TX, CO-ON, FL ("western panhandle"), Cuba
• C. imitatrix Young, TX-IN, VA, FL (ALA, BAK, CHA, DAD, DUV, ESC, HAR, HIG, HOL, LEE, MARI, OSC, PAL, POL, VOL), Cuba
• C. palustris Young, AL, NC, ?FL (DAD), Cuba
• C. slossoni Mutchler, AR-MD-FL (ALA, DAD, PIN), Cuba

Tribe Vatellini

Derovatellus Sharp

• D. lentus floridanus Fall, FL (COLL, DAD, PAL, PAS), Hispaniola, Trinidad

Tribe Hydrovatini

Hydrovatus Motschulsky

• H. inexpectatus Young, FL endemic (ALA, DAD)
• H. peninsularis Young, FL endemic (ALA, HIG, POL)
• H. platycornis Young, GA-FL (ALA, JAC)
• H. pustulatus compressus Sharp, FL (ALA, BRO, CAL, CHA, CIT, COLU, DAD, DUV, GLA, HEN, HIL, LEE, LAK, ORA, OSC, PAL, PIN, POL, SAR, SEM, TAY, WAL), Cuba

Tribe Hyphydrini

Desmopachria Babington

• D. aspera Young, AL?, FL (DAD, PIN, PUT), Bahamas, Cuba
• D. cenchramis Young, FL endemic (BRO, PAL, PAS (first two records from J.H. Epler collection)) (endangered?)
• D. grana LeConte, SC-TX, FL; Young (1954: 56) records the D. grana complex from: FL (ALA, BAY, BRO, DAD, IX, DUV, GLA, HEN, HIL, JEF, LEV, MAD, MARI, MON, PIN, POL, PUT, SAR, TAY, VOL, WAL). However, some species such as D. aspera, D. cenchramis, and D. leechi were subsequently described from Florida. Young (1981: 4) gives the range of the D. grana complex as "Florida" so Roughley has been conservative in assessing county records to species names and has listed only those counties specifically recorded by Young.
• D. leechi Young, SC-MS, FL (ALA, COLL, DAD, FRA, GLA, HIL, LEE, LIB, MON), SAR, TAY, VOL)
• D. mutchleri Blatchley, FL endemic (ALA, BRO, DAD, HER, LEV, PIN)
• D. seminola Young, TX, FL (JAC) (The TX record is based on FSCA specimens found by J.H. Epler)

Pachydrus Sharp

• P. princeps (Blatchley), GA, FL (ALA, BRO, COLU, DAD, FRA, HIG, LAK, LEE, MON, PIN, PAL); According to Young (1953: 115) the type locality on Lake Okeechobee is near the present site of Pahokee, Palm Beach Co., FL.

Tribe Bidessini

Anodocheilus Babington (see Young 1969)

• A. exiguus (Aubé), TX-FL (ALA, BAY, CAL, CLA, DAD, DUV, GUL, HEN, HIL, JAC, LAF, LEO, LEV, LIB, MAD, MARI, OKA, OKE, ORA, OSC, PIN, POL, PUT, SJO, TAY, VOL, WAK, WAL, WAS), Cuba

Bidessonotus Régimbart

• B. longovalis (Blatchley), AL, GA, FL (ALA, GAD, HEN, LEV, MAD, OSC, POL, PUT, TAY, WAK)
• B. inconspicuus (LeConte), OK, MI-NS, FL (ALA, BRE, GAD, JEF, LIB, MAD, SRO)
• B. pulicarius (Aubé), MS, IL-PA, FL (ALA, BRO, CAL, DAD, FLA, GAD, GUL, HAR, HIL, JEF, LEV, LIB, MAD, OKA, PIN, POL, PUT, SJO, SRO, TAY, VOL, WAK, WAL, WAS)

Brachyvatus Zimmermann

• B. apicatus (Clark), FL endemic (ALA, GLA, HIG, MARI, PUT, SAR, VOL), Cuba, Hispaniola, Mexico, CAmer

Liodessus Guignot

• L. affinis (Say), CA, AK-NF, FL (ALA, BRE, FRA, GAD, MARI, MON, TAY, WAK, WAL), Cuba, Bahamas, Mexico, CAmer, SAmer
• L. flavicollis (LeConte), KS, MB-ON, MA, FL (ALA, PUT, WELA)
• L. fuscatus (Crotch), IL, SK-NF, FL (ALA, BAK, BRE, CAL, DUV, GAD, JEFF, MAD, PAL, POL, PUT, TAY)
• L. hobbsi (Young), FL endemic (LIB)

Neobidessus Young

• N. pullus floridanus (Fall), GA, FL (ALA, BRO, CAL, CLA, COLL, DAD, DUV, FLAG, FRA, GLA, HER, HIG, HIL, JEFF, LEE, LEV, MAD, MON, OSC, PAL, PIN, PUT, SAR, SUW, TAY, VOL), Cuba, Bahamas (Andros)
• N. pullus X floridanus, GA, FL (BAY, CAL, GAD, GUL, JAC, WAK, WAL, WAS)
• N. pullus pullus (LeConte), TX, GA, FL (BAY, JAC, SRO, WAL)

Uvarus Guignot

• U. falli (Young), MS, GA, FL (ALA, ?HIL, JEF, LEV, MAD, ?SRO)
• U. granarius (Aubé), IA, MB-NS, FL (ALA, BAK, DUV, GAD, LEV, PIN, PUT, SJO, VOL)
• U. inflatus (Young), NJ, FL (BAY, LIB)
• U. lacustris (Say), KS, ON-VT, FL (CAL, GAD, LIB, MAD, SRO, WAL)
• U. rogersi (Young), GA, FL (BAY, CAL, FRA, GUL, JAC, LEO, LIB, WAK, WAL, WAS)

Tribe Hydroporini

Hydroporus Clairville
n. subg. (Roughley, in litt.)

• H. filiolus ? Fall, MD-FL (GAD)
• H. oblitus ? Aubé, TX, IL-FL (GAD)

subg. Hydroporus (s.str.)

• H. brevicornis Fall, MS, MI-NF, FL (BAY, CAL, GAD, LIB)
• H. dichrous Melsheimer, MO, MN-PQ, ?FL (COLU)
• H. falli Blatchley, AL, FL (ALA, LAK, PIN)
• H. niger Say, TX, ON-NF, FL (BAY, GAD)
• H. ruficeps Aubé, MA-?FL (DUV); See Fall (1923) about this dubious record for Florida.

subg. Neoporus Guignot

• H. asidytus Young, SC, GA, MS, FL (ALA, BAK, FRA, JAC, LIB, WAL)
• H. aulicus Aubé, TX, IN-VA, FL (LEV)
• H. baelus Young, FL endemic (BAY, FRA, LIB, WAK, WAL)
• H. blanchardi Sherman, IN, NY-OK, FL (GAD, HOL)
• H. carolinus complex, IL, ON-NF, FL (ALA, CLA, COLU, HAM, HOL, LIB, MAD, OKA, SJO)
• H. cimicoides Sharp, NJ-FL (ALA, FRA, JAC, LEO, MAD, MARI, PIN, PUT, VOL)
• H. clypealis Sharp, KS, ON-NF, FL (ALA, FLAG, HAM, HIL, LIB, MAD, OKA, OSC, SJO, SRO)
• H. dilatatus Fall, NB-FL (ALA, GAD, HOL, TAY); According to Wolfe (1979) the FL (HOL) record of N. spurius (LeConte) given by Young (1954) is N. dilitatus. Therefore this record is transferred to the latter species.
• H. dixianus Fall, AL, TN-GA, FL (LIB)
• H. effeminatus Fall, MD, AL, FL (BRE, FLAG, LEV, OSC)
• H. floridanus Young, FL endemic (CLA)
• H. gaudens Fall, GA, FL (CAL, GAD, LIB, OKA)
• H. hebes Fall, GA, FL (ALA, BAK, BAY, BRE, FLAG, HIG, HIL, LEV, MAD, MARI, ORA, PIN, POL, PUT, SJO, TAY, VOL); According to Young (1984: 189) the records of H. hebes from Franklin, Liberty and Walton Co. of Young (1954) belong to H. baelus Young and the former species does not occur in the western panhandle of Florida.
• H. helocrinus Young, FL endemic (ALA)
• H. hybridus Aubé, AR, IA-NC, FL (MAD)
• H. lobatus Sharp, AL, MI-NH, FL (ALA, CAL, COLU, GAD, GUL, HAM, LEO, LEV, LIB, MAD, TAY)
• H. lynceus complex, AL, KY-VA, FL (ALA, COLU, FRA, GAD, GUL, LIB, OKA, PUT, SJO, TAY)
• H. mellitus (LeConte), NH-OH-MS-FL (WAL, in J.H. Epler Coll.)
• H. rheocrinus Young, FL endemic (ALA)
• H. shermani Fall, OK, NY-NJ, FL (LIB, GAD)
• H. striatopunctatus Melsheimer, AL, MI-VT, FL (LIB)
• H. uniformis Blatchley, FL endemic (BRO, DAD)
• H. venustus LeConte, AL, IL-NJ, FL (GAD)
• H. vittatipennis Gemminger and Harold, AL, MO-NC, FL (ALA, FLA, HER, LEO, LIB, MAD, SRO, TAY, WAK)

Hygrotus Stephens

• H. berneri Young and Wolfe, FL endemic (DIX, LEO, LIB)
• H. marginipennis (Blatchley), FL endemic (ALA, CHA, DAD, DUV, GLA, HEN, PAL, PIN, SAR, TAY)
• H. nubilus (LeConte), widespread NAmer, FL (JAC, OKA (record from J.H. Epler collection))
• sp. FL (NAS)

Lioporeus Guignot

• L. pilatei Fall, TX-NC, FL (ALA, GIL, LEV)
• L. triangularis (Fall), WI-FL (BAY, record from J.H. Epler collection))

Subfamily Colymbetinae

Tribe Agabini

Agabus Leach

• A. aeruginosus Aubé, MA, IL-AL, FL (COLU, LIB, SJO)
• A. johannis Fall, AL, GA, FL (ALA, BAY, CLA, COLU, HAM, HIL, LIB, ORA, PIN, PUT)
• A. punctatus Melsheimer, MA, MI-IN, FL (ALA)

Ilybius Erichson

• I. oblitus Sharp, MS, KS-NY, FL (ALA, GAD)

Tribe Matini

Matus Aubé

• M. leechi Young, FL endemic (GUL, LIB)
• M. ovatus blatchleyi Leech, AL, GA, FL (ALA, BAY, BRE, DAD, FLAG, FRA, GAD, HIG, HOL, LIB, MAN, OSC, PAL, PIN, SJO, TAY, WAL, WAS)

Tribe Coptotomini

Coptotomus Say: Hilsenhoff (1980) does not list county records for Florida species. Despite a letter to him, Roughley has been unable to get the locality data that he used.

• C. interrogatus Fabricius, MA-AL, FL (35 counties, GLA), Bahamas
• C. lenticus Hilsenhoff, MB, ME-MN, FL (CLA, NAS)
• C. loticus Hilsenhoff, TX, WI-VA, FL (LIB, SRO)
• C. venustus (Say), TX, AR-VA, FL (ALA, CIT, DES, TAY), Mexico

Tribe Colymbetini

Rhantus Stephens

• R. calidus (Fabricius), NY-TX, FL (ALA, BAY, BRO, CIT, CLA, DAD, DUV, FLAG, FRA, GAD, JEFF, LIB, OSC, MON, OSC, PAL, PIN, SJO, TAY, VOL, WAK, WAS), Bahamas, Jamaica, Virgin Islands, Mexico, CAmer, SAmer

Subfamily Dytiscinae

Tribe Eretini

Eretes LaPorte

• E. sticticus (Linnaeus), CA-OK, FL (JAC), Cuba, Mexico

Tribe Hydaticini

Hydaticus Leach

• H. bimarginatus (Say), TX, AR-NY, FL (ALA, BAK, BAY, BRE, BRO, COLU, DAD, DUV, FRA, GAD, GLA, HEN, HIG, HIL, JAC, JEFF, LEE, LEO, LIB, MAN, MON, OKE, ORA, PAL, PIN, POL, PUT, SAR, SEM, SJO, TAY, VOL), Bahamas, Cuba. Young (1954) recorded H. rimosus Aubé from Broward Co., FL, however, Roughley and Pengelly (1982) transferred this record to H. bimarginatus while acknowledging that the former could well be found in Florida as it is presently recorded from Bahamas and Cuba.
• H. cinctipennis Aubé, MA-MS, FL (BAK, WAL)

Tribe Aciliini

Acilius Leach

• A. fraternus dismalus Matta and Michael, VA, NC, FL (ALA, NAS, ?SJO, WAK)
• A. mediatus (Say), MA-FL (SRO)

Graphoderus Dejean

• G. liberus (Say), widespread NAmer, FL (ALA, GAD, LIB)

Thermonectus Dejean

• T. basillaris (Harris), CA, OK-NY, FL (ALA, BAY, BRO, CAL, CHA, COLU, DAD, DUV, FLAG, FRA, GAD, HIG, HIL, JAC, LEV, LIB, MAN, MARI, MON, OKA, OKE, ORA, PAL, POL, PUT, SAR, SEM, SJO, SRO, TAY, WAK, WAL), Bahamas, Cuba, Mexico, Lesser Antilles, SAmer
• T. ornaticollis Aubé, AZ, WY-VT, FL (ALA, BAY, BRO, CAL, CHA, COLU, DAD, DUV, JAC, MART, MON), Bahamas

Tribe Cybistrini

Cybister Curtis

• C. fimbriolatus fimbriolatus (Say), FL endemic (CAL, JAC)
• C. fimbriolatus crotchi Wilke, LA-PA, FL (every county?), Bahamas
• C. occidentalis Aubé, FL (MON), Bahamas, Cuba

Megadytes Sharp

• M. fraternus Sharp, FL (DAD, PAL), Cuba, Bahamas, Jamaica, Hispaniola, Puerto Rico, Lesser Antilles, Guatemala, Panama

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